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While pentose cycle has been long proposed in theory as an intriguing potential glucose metabolism mode with ultra-high NADPH yield, and some carbon recycling at gross flux level has been reported 21,22 , to our knowledge, substantial pentose cycling at net flux level (characterized by reversed GPI net flux, uncoupled glycolysis and oxPPP flux exceeding glucose uptake) has not been experimentally demonstrated in mammalian cells

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